Picker, M.D. & Midgley, J.J. 1996. Pollination by monkey beetles (Coleoptera: Scarabaeidae: Hopliini): flower and colour preferences. African Entomology 4(1): 7–14.
http://pza.sanbi.org/references/picker-md-midgley-jj-1996-pollination-monkey-beetles-coleoptera-scarabaeidae-hopliini